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- Matthew Broderick as Dr. Niko "Nick" Tatopoulos
- Jean Reno as Philippe Roache
- Maria Pitillo as Audrey Timmonds
- Hank Azaria as Victor "Animal" Palotti
- Kevin Dunn as Colonel Hicks
- Michael Lerner as Mayor Ebert
- Lorry Goldman as Gene
- Harry Shearer as Charles Caiman
- Arabella Field as Lucy Palotti
- Vicki Lewis as Dr. Elsie Chapman
- Doug Savant as Sergeant O'Neal
- Malcolm Danare as Dr. Mendel Craven
- Frank Welker as Godzilla (voice)